In the episode Freebird, Rapunzel and Cassandra travel deep into a neighboring forest on their journey and run into Father and Mother. Both characters show that they can turn into birds with magical tea brewed from a magical teapot and can release the transformation with the breaking of a special egg.  It is unclear what the actual relationship is between Father and Mother, but they travel together searching for people that they can change into birds forever.

Mother Bird Form

Mother's bird form

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